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List: DesertLeps
Date: Tue, 24 Jan 2023 12:40:52 -0800 (PST)
From: JONATHAN PELHAM <zapjammer...>
Subject: [NorWestLeps] Taxonomic change - Anthocharis sara complex

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All,

To begin, I don=E2=80=99t assume that everyone in this audience has the sam=
e intense interest in butterfly systematics that I do. However, it is plain=
how important it is to Todd Stout, Harry Pavulaan & Steve Kohler, all enga=
ged in preparing regional treatments. Further, Mike Fisher, Jim Brock and K=
en Davenport have long been familiar with issues of butterfly systematics a=
s they prepared their work.

Me? I catalogue it all and have a commitment to it=E2=80=99s users. As resu=
lt, I am familiar with what has transpired and amazed at what is upon us cu=
rrently. Apparently, no matter how elusive a definitive systematic arrangem=
ent of butterflies may be, it can be attained. Perhaps I will witness it!

It may be useful to detail how past taxonomic changes have taken place. It =
gives me pleasure to document one example reflecting the hard work of one o=
f our own - Todd Stout- and the reticence of this cataloguer to make BIG ch=
anges without profound support.

In 1986 Geiger & Shapiro published a paper giving electrophoretic evidence =
supporting Anthocharis sara, A.stella and A.julia as =E2=80=9Cdifferentiate=
d at the species level.=E2=80=9D

As I recall, this was discussed but not acted upon until Layberry et al tre=
ated all Canadian populations as A.stella in 1998. A year later, Opler doub=
led down, adding a twist, treating A. thoosa as a distinct species as well.

These publications created difficulties as I prepared the catalogue but I h=
ad a friend (TS) who had published a paper in 2005 treating A.sara and A.st=
ella as distinct species in a rearing study. We talked and he said (somethi=
ng like): JP...I=E2=80=99m crazy for orange-tips and I=E2=80=99m going to f=
igure this one out - but I haven=E2=80=99t yet.=E2=80=9D

I deferred from accepting the split in 2008. I eagerly awaited Todd=E2=80=
=99s paper...which arrived in 2018 and I am gratified that Opler et al, 202=
2 supported Todd=E2=80=99s conclusions

This took 22 years but I am not citing this as any sort of standard! It is =
only an example.

JPPelham


Geiger, Hansj=C3=BCrg and Arthur Maurice Shapiro
1986 Electrophoretic evidence for speciation within the nominal species Ant=
hocharis sara Lucas (Pieridae). Journal of Research on the Lepidoptera 25(1=
): 15-24, 2 figs., 4 tbls. {Spring 1986, 1 Oct 1986}

Layberry, Ross A., Peter W. Hall and James Donald Lafontaine (plates by Joh=
n T. Fowler)
1998 The butterflies of Canada. Toronto, Buffalo & London; University of To=
ronto Press: 280 pp., 32 pls., 294 maps and ill. {1998}


Opler, Paul Alexander
1999 A field guide to western butterflies. Boston, New York; Houghton Miffl=
in Company, The Peterson Field Guide Series (Amy Bartlett Wright (illustrat=
or): xiv, [2], 540 pp., 44 pls., figs. & maps. {1999}

Stout, Todd L.
2005 The challenge of raising northern Utah orangetips (Anthocharis stella =
browningi and Anthocharis sara thoosa). Bulletin of the Utah Lepidopterists=
=E2=80=99 Society 12(1): 5-9, 13, 7 figs. {Mar 2005}

Scott, James Allen and Michael S. Fisher
2008 Anthocharis =E2=80=9Csara=E2=80=9D group, especially in Colorado and v=
icinity (with new research from Todd Stout) (one ssp. coauthored by Norbert=
G. Kondla). Pp. 1-14, pl. 2, in: J. Scott (Ed.), Geographic variation and =
new taxa of western North American butterflies, especially from Colorado. P=
apilio (New Series) 18 {3 Dec 2008}

Stout, Todd L.
2010 Observations on Anthocharis julia browningi and Anthocharis thoosa tho=
osa including tension zones near Nephi, Juab County, Utah. Taxonomic Report=
of the International Lepidoptera Survey 7(4): 1-12 {31 May 2010}

Stout, Todd L.
2018 A review of three species-level taxa of the Anthocharis sara complex (=
Lepidoptera: Pieridae: Pierinae: Anthocharidini). Insecta Mundi 0615: 1-38,=
26 figs., 3 tbls.,1 app. {30 Mar 2018}

Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Qian Cong. J=
inhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin
2022 DNA barcodes reveal different speciation scenarios in the four North A=
merican Anthocharis Boisduval, Rambur, [Dum=C3=A9nil] & Graslin, [1833] (Le=
pidoptera: Pieridae: Pierinae: Anthocharidini) species groups. Zootaxa 5194=
(4): 519=E2=80=93539, 28 figs. {7 Oct 2022}


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<html>
<head>=20
<meta charset=3D"UTF-8">=20
</head>
<body>
<div>
<br>
</div>
<div>
All,
</div>
<div>
<br>
</div>
<div>
To begin, I don=E2=80=99t assume that everyone in this audience has the =
same intense interest in butterfly systematics that I do. However, it is pl=
ain how important it is to Todd Stout, Harry Pavulaan &amp; Steve Kohler, a=
ll engaged in preparing regional treatments. Further, Mike Fisher, Jim Broc=
k and Ken Davenport have long been familiar with issues of butterfly system=
atics as they prepared their work.
</div>
<div>
<br>
</div>
<div>
Me? I catalogue it all and have a commitment to it=E2=80=99s users. As r=
esult, I am familiar with what has transpired and amazed at what is upon us=
currently. Apparently, no matter how elusive a definitive systematic arran=
gement of butterflies may be, it can be attained. Perhaps I will witness it=
!
</div>
<div>
<br>
</div>
<div>
It may be useful to detail how past taxonomic changes have taken place. =
It gives me pleasure to document one example reflecting the hard work of on=
e of our own - Todd Stout- and the reticence of this cataloguer to make BIG=
changes without profound support.
</div>
<div>
<br>
</div>
<div>
In 1986 Geiger &amp; Shapiro published a paper giving electrophoretic ev=
idence supporting Anthocharis sara, A.stella and A.julia as =E2=80=9Cdiffer=
entiated at the species level.=E2=80=9D
</div>
<div>
<br>
</div>
<div>
As I recall, this was discussed but not acted upon until Layberry et al =
treated all Canadian populations as A.stella in 1998. A year later, Opler d=
oubled down, adding a twist, treating A. thoosa as a distinct species as we=
ll.
</div>
<div>
<br>
</div>
<div>
These publications created difficulties as I prepared the catalogue but =
I had a friend (TS) who had published a paper in 2005 treating A.sara and A=
.stella as distinct species in a rearing study. We talked and he said (some=
thing like): JP...I=E2=80=99m crazy for orange-tips and I=E2=80=99m going t=
o figure this one out - but I haven=E2=80=99t yet.=E2=80=9D
</div>
<div>
<br>
</div>
<div>
I deferred from accepting the split in 2008. I eagerly awaited Todd=E2=
=80=99s paper...which arrived in 2018 and I am gratified that Opler et al, =
2022 supported Todd=E2=80=99s conclusions
</div>
<div>
<br>
</div>
<div>
This took 22 years but I am not citing this as any sort of standard! It =
is only an example.
</div>
<div>
<br>
</div>
<div>
JPPelham
</div>
<div>
<br>
</div>
<div>
<br>
</div>
<div>
<strong>Geiger, Hansj=C3=BCrg and Arthur Maurice Shapiro</strong>
</div>
<div>
1986 Electrophoretic evidence for speciation within the nominal species =
<em>Anthocharis sara</em> Lucas (Pieridae). <em>Journal of Research on the =
Lepidoptera 25(1): 15-24, 2 figs., 4 tbls. {Spring 1986, 1 Oct 1986}</em>
</div>
<div>
<br>
</div>
<div>
<strong>Layberry, Ross A., Peter W. Hall and James Donald Lafontaine</st=
rong> (plates by John T. Fowler)
</div>
<div>
1998 <em>The butterflies of Canada</em>. Toronto, Buffalo &amp; London; =
University of Toronto Press: 280 pp., 32 pls., 294 maps and ill. {1998}
</div>
<div>
<br>
</div>
<div>
<br>
</div>
<div>
<strong>Opler, Paul Alexander</strong>
</div>
<div>
1999 <em>A field guide to western butterflies</em>. Boston, New York; Ho=
ughton Mifflin Company, The Peterson Field Guide Series (Amy Bartlett Wrigh=
t (illustrator): xiv, [2], 540 pp., 44 pls., figs. &amp; maps. {1999}
</div>
<div>
<br>
</div>
<div>
<strong>Stout, Todd L.</strong>
</div>
<div>
2005 The challenge of raising northern Utah orangetips (<em>Anthocharis =
stella browningi</em> and <em>Anthocharis sara thoosa</em>). <em>Bulletin o=
f the Utah Lepidopterists=E2=80=99 Society 12(1): 5-9, 13, 7 figs. {Mar 200=
5}</em>
</div>
<div style=3D"font-size: 12pt; font-family: helvetica, arial, sans-serif;=
color: rgb(51, 51, 51);" class=3D"default-style">
<br>
</div>
<div>
<strong>Scott, James Allen and Michael S. Fisher</strong>
</div>
<div>
2008 <em>Anthocharis</em> =E2=80=9C<em>sara</em>=E2=80=9D group, especia=
lly in Colorado and vicinity (with new research from Todd Stout) (one ssp. =
coauthored by Norbert G. Kondla). Pp. 1-14, pl. 2, in: J. Scott (Ed.), Geog=
raphic variation and new taxa of western North American butterflies, especi=
ally from Colorado. <em>Papilio (New Series) 18 {3 Dec 2008}</em>
</div>
<div>
<br>
</div>
<div>
<strong>Stout, Todd L.</strong>
</div>
<div>
2010 Observations on <em>Anthocharis julia browningi</em> and <em>Anthoc=
haris thoosa thoosa</em> including tension zones near Nephi, Juab County, U=
tah. <em>Taxonomic Report of the International Lepidoptera Survey 7(4): 1-1=
2 {31 May 2010}</em>
</div>
<div>
<br>
</div>
<div>
<strong>Stout, Todd L.</strong>
</div>
<div>
2018 A review of three species-level taxa of the <em>Anthocharis sara</e=
m> complex (Lepidoptera: Pieridae: Pierinae: Anthocharidini). <em>Insecta M=
undi 0615: 1-38, 26 figs., 3 tbls.,1 app. {30 Mar 2018}</em>
</div>
<div>
<br>
</div>
<div>
<strong>Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Q=
ian Cong. Jinhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin</strong>
</div>
<div>
2022 DNA barcodes reveal different speciation scenarios in the four Nort=
h American <em>Anthocharis</em> Boisduval, Rambur, [Dum=C3=A9nil] &amp; Gra=
slin, [1833] (Lepidoptera: Pieridae: Pierinae: Anthocharidini) species grou=
ps. <em>Zootaxa 5194(4): 519=E2=80=93539, 28 figs. {7 Oct 2022}</em>
</div>
</body>
</html>


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