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| List: DesertLeps Date: Tue, 24 Jan 2023 12:40:52 -0800 (PST) From: JONATHAN PELHAM <zapjammer...> Subject: [NorWestLeps] Taxonomic change - Anthocharis sara complex |
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------=_Part_1937613_252156105.1674592852666 MIME-Version: 1.0 Content-Type: text/plain; charset=UTF-8 Content-Transfer-Encoding: quoted-printable All, To begin, I don=E2=80=99t assume that everyone in this audience has the sam= e intense interest in butterfly systematics that I do. However, it is plain= how important it is to Todd Stout, Harry Pavulaan & Steve Kohler, all enga= ged in preparing regional treatments. Further, Mike Fisher, Jim Brock and K= en Davenport have long been familiar with issues of butterfly systematics a= s they prepared their work. Me? I catalogue it all and have a commitment to it=E2=80=99s users. As resu= lt, I am familiar with what has transpired and amazed at what is upon us cu= rrently. Apparently, no matter how elusive a definitive systematic arrangem= ent of butterflies may be, it can be attained. Perhaps I will witness it! It may be useful to detail how past taxonomic changes have taken place. It = gives me pleasure to document one example reflecting the hard work of one o= f our own - Todd Stout- and the reticence of this cataloguer to make BIG ch= anges without profound support. In 1986 Geiger & Shapiro published a paper giving electrophoretic evidence = supporting Anthocharis sara, A.stella and A.julia as =E2=80=9Cdifferentiate= d at the species level.=E2=80=9D As I recall, this was discussed but not acted upon until Layberry et al tre= ated all Canadian populations as A.stella in 1998. A year later, Opler doub= led down, adding a twist, treating A. thoosa as a distinct species as well. These publications created difficulties as I prepared the catalogue but I h= ad a friend (TS) who had published a paper in 2005 treating A.sara and A.st= ella as distinct species in a rearing study. We talked and he said (somethi= ng like): JP...I=E2=80=99m crazy for orange-tips and I=E2=80=99m going to f= igure this one out - but I haven=E2=80=99t yet.=E2=80=9D I deferred from accepting the split in 2008. I eagerly awaited Todd=E2=80= =99s paper...which arrived in 2018 and I am gratified that Opler et al, 202= 2 supported Todd=E2=80=99s conclusions This took 22 years but I am not citing this as any sort of standard! It is = only an example. JPPelham Geiger, Hansj=C3=BCrg and Arthur Maurice Shapiro 1986 Electrophoretic evidence for speciation within the nominal species Ant= hocharis sara Lucas (Pieridae). Journal of Research on the Lepidoptera 25(1= ): 15-24, 2 figs., 4 tbls. {Spring 1986, 1 Oct 1986} Layberry, Ross A., Peter W. Hall and James Donald Lafontaine (plates by Joh= n T. Fowler) 1998 The butterflies of Canada. Toronto, Buffalo & London; University of To= ronto Press: 280 pp., 32 pls., 294 maps and ill. {1998} Opler, Paul Alexander 1999 A field guide to western butterflies. Boston, New York; Houghton Miffl= in Company, The Peterson Field Guide Series (Amy Bartlett Wright (illustrat= or): xiv, [2], 540 pp., 44 pls., figs. & maps. {1999} Stout, Todd L. 2005 The challenge of raising northern Utah orangetips (Anthocharis stella = browningi and Anthocharis sara thoosa). Bulletin of the Utah Lepidopterists= =E2=80=99 Society 12(1): 5-9, 13, 7 figs. {Mar 2005} Scott, James Allen and Michael S. Fisher 2008 Anthocharis =E2=80=9Csara=E2=80=9D group, especially in Colorado and v= icinity (with new research from Todd Stout) (one ssp. coauthored by Norbert= G. Kondla). Pp. 1-14, pl. 2, in: J. Scott (Ed.), Geographic variation and = new taxa of western North American butterflies, especially from Colorado. P= apilio (New Series) 18 {3 Dec 2008} Stout, Todd L. 2010 Observations on Anthocharis julia browningi and Anthocharis thoosa tho= osa including tension zones near Nephi, Juab County, Utah. Taxonomic Report= of the International Lepidoptera Survey 7(4): 1-12 {31 May 2010} Stout, Todd L. 2018 A review of three species-level taxa of the Anthocharis sara complex (= Lepidoptera: Pieridae: Pierinae: Anthocharidini). Insecta Mundi 0615: 1-38,= 26 figs., 3 tbls.,1 app. {30 Mar 2018} Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Qian Cong. J= inhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin 2022 DNA barcodes reveal different speciation scenarios in the four North A= merican Anthocharis Boisduval, Rambur, [Dum=C3=A9nil] & Graslin, [1833] (Le= pidoptera: Pieridae: Pierinae: Anthocharidini) species groups. Zootaxa 5194= (4): 519=E2=80=93539, 28 figs. {7 Oct 2022} -=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D- Groups.io Links: You receive all messages sent to this group. View/Reply Online (#11201): https://groups.io/g/NorWestLeps/message/11201 Mute This Topic: https://groups.io/mt/96507103/795800 Group Owner: NorWestLeps+<owner...> Unsubscribe: https://groups.io/g/NorWestLeps/leave/4559582/795800/778100922= /xyzzy [<lists...>] -=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D- ------=_Part_1937613_252156105.1674592852666 MIME-Version: 1.0 Content-Type: text/html; charset=UTF-8 Content-Transfer-Encoding: quoted-printable <!doctype html> <html> <head>=20 <meta charset=3D"UTF-8">=20 </head> <body> <div> <br> </div> <div> All, </div> <div> <br> </div> <div> To begin, I don=E2=80=99t assume that everyone in this audience has the = same intense interest in butterfly systematics that I do. However, it is pl= ain how important it is to Todd Stout, Harry Pavulaan & Steve Kohler, a= ll engaged in preparing regional treatments. Further, Mike Fisher, Jim Broc= k and Ken Davenport have long been familiar with issues of butterfly system= atics as they prepared their work. </div> <div> <br> </div> <div> Me? I catalogue it all and have a commitment to it=E2=80=99s users. As r= esult, I am familiar with what has transpired and amazed at what is upon us= currently. Apparently, no matter how elusive a definitive systematic arran= gement of butterflies may be, it can be attained. Perhaps I will witness it= ! </div> <div> <br> </div> <div> It may be useful to detail how past taxonomic changes have taken place. = It gives me pleasure to document one example reflecting the hard work of on= e of our own - Todd Stout- and the reticence of this cataloguer to make BIG= changes without profound support. </div> <div> <br> </div> <div> In 1986 Geiger & Shapiro published a paper giving electrophoretic ev= idence supporting Anthocharis sara, A.stella and A.julia as =E2=80=9Cdiffer= entiated at the species level.=E2=80=9D </div> <div> <br> </div> <div> As I recall, this was discussed but not acted upon until Layberry et al = treated all Canadian populations as A.stella in 1998. A year later, Opler d= oubled down, adding a twist, treating A. thoosa as a distinct species as we= ll. </div> <div> <br> </div> <div> These publications created difficulties as I prepared the catalogue but = I had a friend (TS) who had published a paper in 2005 treating A.sara and A= .stella as distinct species in a rearing study. We talked and he said (some= thing like): JP...I=E2=80=99m crazy for orange-tips and I=E2=80=99m going t= o figure this one out - but I haven=E2=80=99t yet.=E2=80=9D </div> <div> <br> </div> <div> I deferred from accepting the split in 2008. I eagerly awaited Todd=E2= =80=99s paper...which arrived in 2018 and I am gratified that Opler et al, = 2022 supported Todd=E2=80=99s conclusions </div> <div> <br> </div> <div> This took 22 years but I am not citing this as any sort of standard! It = is only an example. </div> <div> <br> </div> <div> JPPelham </div> <div> <br> </div> <div> <br> </div> <div> <strong>Geiger, Hansj=C3=BCrg and Arthur Maurice Shapiro</strong> </div> <div> 1986 Electrophoretic evidence for speciation within the nominal species = <em>Anthocharis sara</em> Lucas (Pieridae). <em>Journal of Research on the = Lepidoptera 25(1): 15-24, 2 figs., 4 tbls. {Spring 1986, 1 Oct 1986}</em> </div> <div> <br> </div> <div> <strong>Layberry, Ross A., Peter W. Hall and James Donald Lafontaine</st= rong> (plates by John T. Fowler) </div> <div> 1998 <em>The butterflies of Canada</em>. Toronto, Buffalo & London; = University of Toronto Press: 280 pp., 32 pls., 294 maps and ill. {1998} </div> <div> <br> </div> <div> <br> </div> <div> <strong>Opler, Paul Alexander</strong> </div> <div> 1999 <em>A field guide to western butterflies</em>. Boston, New York; Ho= ughton Mifflin Company, The Peterson Field Guide Series (Amy Bartlett Wrigh= t (illustrator): xiv, [2], 540 pp., 44 pls., figs. & maps. {1999} </div> <div> <br> </div> <div> <strong>Stout, Todd L.</strong> </div> <div> 2005 The challenge of raising northern Utah orangetips (<em>Anthocharis = stella browningi</em> and <em>Anthocharis sara thoosa</em>). <em>Bulletin o= f the Utah Lepidopterists=E2=80=99 Society 12(1): 5-9, 13, 7 figs. {Mar 200= 5}</em> </div> <div style=3D"font-size: 12pt; font-family: helvetica, arial, sans-serif;= color: rgb(51, 51, 51);" class=3D"default-style"> <br> </div> <div> <strong>Scott, James Allen and Michael S. Fisher</strong> </div> <div> 2008 <em>Anthocharis</em> =E2=80=9C<em>sara</em>=E2=80=9D group, especia= lly in Colorado and vicinity (with new research from Todd Stout) (one ssp. = coauthored by Norbert G. Kondla). Pp. 1-14, pl. 2, in: J. Scott (Ed.), Geog= raphic variation and new taxa of western North American butterflies, especi= ally from Colorado. <em>Papilio (New Series) 18 {3 Dec 2008}</em> </div> <div> <br> </div> <div> <strong>Stout, Todd L.</strong> </div> <div> 2010 Observations on <em>Anthocharis julia browningi</em> and <em>Anthoc= haris thoosa thoosa</em> including tension zones near Nephi, Juab County, U= tah. <em>Taxonomic Report of the International Lepidoptera Survey 7(4): 1-1= 2 {31 May 2010}</em> </div> <div> <br> </div> <div> <strong>Stout, Todd L.</strong> </div> <div> 2018 A review of three species-level taxa of the <em>Anthocharis sara</e= m> complex (Lepidoptera: Pieridae: Pierinae: Anthocharidini). <em>Insecta M= undi 0615: 1-38, 26 figs., 3 tbls.,1 app. {30 Mar 2018}</em> </div> <div> <br> </div> <div> <strong>Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Q= ian Cong. Jinhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin</strong> </div> <div> 2022 DNA barcodes reveal different speciation scenarios in the four Nort= h American <em>Anthocharis</em> Boisduval, Rambur, [Dum=C3=A9nil] & Gra= slin, [1833] (Lepidoptera: Pieridae: Pierinae: Anthocharidini) species grou= ps. <em>Zootaxa 5194(4): 519=E2=80=93539, 28 figs. {7 Oct 2022}</em> </div> </body> </html> <div width=3D"1" style=3D"color:white;clear:both">_._,_._,_</div> <hr> Groups.io Links:<p> =20 You receive all messages sent to this group. =20 =20 <p> <a target=3D"_blank" href=3D"https://groups.io/g/NorWestLeps/message/11201"= >View/Reply Online (#11201)</a> | <a target=3D"_blank" href=3D"mailto:<NorWestLeps...>?subject=3DRe:%2= 0%5BNorWestLeps%5D%20Taxonomic%20change%20-%20Anthocharis%20sara%20complex"= >Reply To Group</a> =20 | <a target=3D"_blank" href=3D"mailto:<zapjammer...>?subject=3DP= rivate:%20Re:%20%5BNorWestLeps%5D%20Taxonomic%20change%20-%20Anthocharis%20= sara%20complex">Reply To Sender</a> =20 | <a target=3D"_blank" href=3D"https://groups.io/mt/96507103/795800">Mute T= his Topic</a> | <a href=3D"https://groups.io/g/NorWestLeps/post">New Topic</a><br> <a href=3D"https://groups.io/g/NorWestLeps/editsub/795800">Your Subscriptio= n</a> | <a href=3D"mailto:NorWestLeps+<owner...>">Contact Group Owner</a> | <a href=3D"https://groups.io/g/NorWestLeps/leave/4559582/795800/778100922/x= yzzy">Unsubscribe</a> [<lists...>]<br> <div width=3D"1" style=3D"color:white;clear:both">_._,_._,_</div> ------=_Part_1937613_252156105.1674592852666-- |
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