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| List: NorWest Leps Date: Tue, 24 Jan 2023 14:18:33 -0800 From: Neil Bjorklund <nel-de-bearck...> Subject: Re: [NorWestLeps] Taxonomic change - Anthocharis sara complex |
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--B_3757414716_2384768163 Content-type: text/plain; charset="UTF-8" Content-transfer-encoding: quoted-printable Thanks for sharing this Jon!=C2=A0 I read the 2022 paper with interest.=C2= =A0 Then I groaned when I saw Figure 23, the map for the A. sara group.=C2= =A0 As I read it, it documents A. sara in the northern part of the coast ra= nge of Oregon, likely flying in sympatry with A. julia, and probably occurs= throughout the coast range.=C2=A0 Which means lazy amatuer field lepidopte= rists like me can=E2=80=99t rely solely on a simple geographic forumla to s= eparate the two species (e.g., =E2=80=9Cif its north of Jackson and Josephi= ne counties, it A. julia=E2=80=9D). And photos may be of no help in determi= ning the ID. As the paper states, =E2=80=9CThis is problematic because over= lap in adult morphological characters of all three species make it challeng= ing to identify wing traits exclusive to any single species.=E2=80=9D Groan= .=C2=A0 And then, =E2=80=9CThese similar phenotypes are distributed randoml= y throughout the sara group taxa and are not regionally correlated except f= or those which are phenotypically similar and fly in near sympatry=E2=80=A6= =E2=80=9D=C2=A0 I interpret the latter to mean that one would have to learn= the wing trait differences of the two species within the areas in which th= ey fly in sympatry.=C2=A0 Groan. =20 Someone please let me know if I=E2=80=99m misinterpreting this=E2=80=A6 =20 Thanks, =20 Neil Bjorklund www.butterfliesoforegon.com=20 =20 =20 =20 From: <NorWestLeps...> on behalf of JONATHAN PELHAM <zapjammer@comca= st.net> Reply-To: <NorWestLeps...> Date: Tuesday, January 24, 2023 at 12:40 PM To: <DesertLeps...> Cc: TILS <TILS...>, desertleps <desertleps...>, norwestleps <= <norwestleps...> Subject: [NorWestLeps] Taxonomic change - Anthocharis sara complex =20 =20 All,=20 =20 To begin, I don=E2=80=99t assume that everyone in this audience has the sam= e intense interest in butterfly systematics that I do. However, it is plain= how important it is to Todd Stout, Harry Pavulaan & Steve Kohler, all enga= ged in preparing regional treatments. Further, Mike Fisher, Jim Brock and K= en Davenport have long been familiar with issues of butterfly systematics a= s they prepared their work.=20 =20 Me? I catalogue it all and have a commitment to it=E2=80=99s users. As resu= lt, I am familiar with what has transpired and amazed at what is upon us cu= rrently. Apparently, no matter how elusive a definitive systematic arrangem= ent of butterflies may be, it can be attained. Perhaps I will witness it!= =20 =20 It may be useful to detail how past taxonomic changes have taken place. It = gives me pleasure to document one example reflecting the hard work of one o= f our own - Todd Stout- and the reticence of this cataloguer to make BIG ch= anges without profound support.=20 =20 In 1986 Geiger & Shapiro published a paper giving electrophoretic evidence = supporting Anthocharis sara, A.stella and A.julia as =E2=80=9Cdifferentiate= d at the species level.=E2=80=9D=20 =20 As I recall, this was discussed but not acted upon until Layberry et al tre= ated all Canadian populations as A.stella in 1998. A year later, Opler doub= led down, adding a twist, treating A. thoosa as a distinct species as well.= =20 =20 These publications created difficulties as I prepared the catalogue but I h= ad a friend (TS) who had published a paper in 2005 treating A.sara and A.st= ella as distinct species in a rearing study. We talked and he said (somethi= ng like): JP...I=E2=80=99m crazy for orange-tips and I=E2=80=99m going to f= igure this one out - but I haven=E2=80=99t yet.=E2=80=9D=20 =20 I deferred from accepting the split in 2008. I eagerly awaited Todd=E2=80= =99s paper...which arrived in 2018 and I am gratified that Opler et al, 202= 2 supported Todd=E2=80=99s conclusions=20 =20 This took 22 years but I am not citing this as any sort of standard! It is = only an example.=20 =20 JPPelham=20 =20 =20 Geiger, Hansj=C3=BCrg and Arthur Maurice Shapiro=20 1986 Electrophoretic evidence for speciation within the nominal species Ant= hocharis sara Lucas (Pieridae). Journal of Research on the Lepidoptera 25(1= ): 15-24, 2 figs., 4 tbls. {Spring 1986, 1 Oct 1986}=20 =20 Layberry, Ross A., Peter W. Hall and James Donald Lafontaine (plates by Joh= n T. Fowler)=20 1998 The butterflies of Canada. Toronto, Buffalo & London; University of To= ronto Press: 280 pp., 32 pls., 294 maps and ill. {1998}=20 =20 =20 Opler, Paul Alexander=20 1999 A field guide to western butterflies. Boston, New York; Houghton Miffl= in Company, The Peterson Field Guide Series (Amy Bartlett Wright (illustrat= or): xiv, [2], 540 pp., 44 pls., figs. & maps. {1999}=20 =20 Stout, Todd L.=20 2005 The challenge of raising northern Utah orangetips (Anthocharis stella = browningi and Anthocharis sara thoosa). Bulletin of the Utah Lepidopterists= =E2=80=99 Society 12(1): 5-9, 13, 7 figs. {Mar 2005}=20 =20 Scott, James Allen and Michael S. Fisher=20 2008 Anthocharis =E2=80=9Csara=E2=80=9D group, especially in Colorado and v= icinity (with new research from Todd Stout) (one ssp. coauthored by Norbert= G. Kondla). Pp. 1-14, pl. 2, in: J. Scott (Ed.), Geographic variation and = new taxa of western North American butterflies, especially from Colorado. P= apilio (New Series) 18 {3 Dec 2008}=20 =20 Stout, Todd L.=20 2010 Observations on Anthocharis julia browningi and Anthocharis thoosa tho= osa including tension zones near Nephi, Juab County, Utah. Taxonomic Report= of the International Lepidoptera Survey 7(4): 1-12 {31 May 2010}=20 =20 Stout, Todd L.=20 2018 A review of three species-level taxa of the Anthocharis sara complex (= Lepidoptera: Pieridae: Pierinae: Anthocharidini). Insecta Mundi 0615: 1-38,= 26 figs., 3 tbls.,1 app. {30 Mar 2018}=20 =20 Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Qian Cong. J= inhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin=20 2022 DNA barcodes reveal different speciation scenarios in the four North A= merican Anthocharis Boisduval, Rambur, [Dum=C3=A9nil] & Graslin, [1833] (Le= pidoptera: Pieridae: Pierinae: Anthocharidini) species groups. Zootaxa 5194= (4): 519=E2=80=93539, 28 figs. {7 Oct 2022}=20 -=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D- Groups.io Links: You receive all messages sent to this group. View/Reply Online (#11202): https://groups.io/g/NorWestLeps/message/11202 Mute This Topic: https://groups.io/mt/96507103/795800 Group Owner: NorWestLeps+<owner...> Unsubscribe: https://groups.io/g/NorWestLeps/leave/4559582/795800/778100922= /xyzzy [<lists...>] -=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D-=3D- --B_3757414716_2384768163 Content-type: text/html; charset="UTF-8" Content-transfer-encoding: quoted-printable <html xmlns:v=3D"urn:schemas-microsoft-com:vml" xmlns:o=3D"urn:schemas-micr= osoft-com:office:office" xmlns:w=3D"urn:schemas-microsoft-com:office:word" = xmlns:m=3D"http://schemas.microsoft.com/office/2004/12/omml" xmlns=3D"http:= //www.w3.org/TR/REC-html40"><head><meta http-equiv=3DContent-Type content= =3D"text/html; charset=3Dutf-8"><meta name=3DGenerator content=3D"Microsoft= Word 15 (filtered medium)"><!--[if !mso]><style>v\:* {behavior:url(#defaul= t#VML);} o\:* {behavior:url(#default#VML);} w\:* {behavior:url(#default#VML);} .shape {behavior:url(#default#VML);} </style><![endif]--><style><!-- /* Font Definitions */ @font-face {font-family:Helvetica; panose-1:0 0 0 0 0 0 0 0 0 0;} @font-face {font-family:"Cambria Math"; panose-1:2 4 5 3 5 4 6 3 2 4;} @font-face {font-family:Calibri; panose-1:2 15 5 2 2 2 4 3 2 4;} /* Style Definitions */ p.MsoNormal, li.MsoNormal, div.MsoNormal {margin:0in; font-size:11.0pt; font-family:"Calibri",sans-serif;} a:link, span.MsoHyperlink {mso-style-priority:99; color:blue; text-decoration:underline;} span.EmailStyle21 {mso-style-type:personal-reply; font-family:"Calibri",sans-serif; color:windowtext;} .MsoChpDefault {mso-style-type:export-only; font-size:10.0pt;} @page WordSection1 {size:8.5in 11.0in; margin:1.0in 1.0in 1.0in 1.0in;} div.WordSection1 {page:WordSection1;} --></style><!--[if gte mso 9]><xml> <o:shapedefaults v:ext=3D"edit" spidmax=3D"1026" /> </xml><![endif]--><!--[if gte mso 9]><xml> <o:shapelayout v:ext=3D"edit"> <o:idmap v:ext=3D"edit" data=3D"1" /> </o:shapelayout></xml><![endif]--></head><body lang=3DEN-US link=3Dblue vli= nk=3Dpurple style=3D'word-wrap:break-word'><div class=3DWordSection1><p cla= ss=3DMsoNormal>Thanks for sharing this Jon!=C2=A0 I read the 2022 paper wit= h interest.=C2=A0 Then I groaned when I saw Figure 23, the map for the <i>A= . sara</i> group.=C2=A0 As I read it, it documents <i>A. sara</i> in the no= rthern part of the coast range of Oregon, likely flying in sympatry with <i= >A. julia, and probably occurs throughout the coast range.=C2=A0 </i>Which = means lazy amatuer field lepidopterists like me can=E2=80=99t rely solely o= n a simple geographic forumla to separate the two species (e.g., =E2=80=9Ci= f its north of Jackson and Josephine counties, it <i>A. julia</i>=E2=80=9D)= . And photos may be of no help in determining the ID. As the paper states, = =E2=80=9CThis is problematic because overlap in adult morphological charact= ers of all three species make it challenging to identify wing traits exclus= ive to any single species.=E2=80=9D Groan.=C2=A0 And then, =E2=80=9CThese s= imilar phenotypes are distributed randomly throughout the sara group taxa a= nd are not regionally correlated except for those which are phenotypically = similar and fly in near sympatry=E2=80=A6=E2=80=9D=C2=A0 I interpret the la= tter to mean that one would have to learn the wing trait differences of the= two species <i>within the areas in which they fly in sympatry</i>.=C2=A0 G= roan.<o:p></o:p></p><p class=3DMsoNormal><o:p> </o:p></p><p class=3DMs= oNormal>Someone please let me know if I=E2=80=99m misinterpreting this=E2= =80=A6<o:p></o:p></p><p class=3DMsoNormal><o:p> </o:p></p><p class=3DM= soNormal>Thanks,<o:p></o:p></p><p class=3DMsoNormal><o:p> </o:p></p><p= class=3DMsoNormal>Neil Bjorklund<o:p></o:p></p><p class=3DMsoNormal><a hre= f=3D"http://www.butterfliesoforegon.com">www.butterfliesoforegon.com</a> <o= :p></o:p></p><p class=3DMsoNormal><o:p> </o:p></p><p class=3DMsoNormal= ><o:p> </o:p></p><p class=3DMsoNormal><o:p> </o:p></p><div style= =3D'border:none;border-top:solid #B5C4DF 1.0pt;padding:3.0pt 0in 0in 0in'><= p class=3DMsoNormal><b><span style=3D'font-size:12.0pt;color:black'>From: <= /span></b><span style=3D'font-size:12.0pt;color:black'><NorWestLeps@grou= ps.io> on behalf of JONATHAN PELHAM <<zapjammer...>><br><b>= Reply-To: </b><<NorWestLeps...>><br><b>Date: </b>Tuesday, Januar= y 24, 2023 at 12:40 PM<br><b>To: </b><<DesertLeps...>>= ;<br><b>Cc: </b>TILS <<TILS...>>, desertleps <desertleps@grou= ps.io>, norwestleps <<norwestleps...>><br><b>Subject: </b>[No= rWestLeps] Taxonomic change - Anthocharis sara complex<o:p></o:p></span></p= ></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div><p class= =3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNormal>All, <o:p= ></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div>= <p class=3DMsoNormal>To begin, I don=E2=80=99t assume that everyone in this= audience has the same intense interest in butterfly systematics that I do.= However, it is plain how important it is to Todd Stout, Harry Pavulaan &am= p; Steve Kohler, all engaged in preparing regional treatments. Further, Mik= e Fisher, Jim Brock and Ken Davenport have long been familiar with issues o= f butterfly systematics as they prepared their work. <o:p></o:p></p></div><= div><p class=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNorm= al>Me? I catalogue it all and have a commitment to it=E2=80=99s users. As r= esult, I am familiar with what has transpired and amazed at what is upon us= currently. Apparently, no matter how elusive a definitive systematic arran= gement of butterflies may be, it can be attained. Perhaps I will witness it= ! <o:p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div= ><div><p class=3DMsoNormal>It may be useful to detail how past taxonomic ch= anges have taken place. It gives me pleasure to document one example reflec= ting the hard work of one of our own - Todd Stout- and the reticence of thi= s cataloguer to make BIG changes without profound support. <o:p></o:p></p><= /div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DM= soNormal>In 1986 Geiger & Shapiro published a paper giving electrophore= tic evidence supporting Anthocharis sara, A.stella and A.julia as =E2=80=9C= differentiated at the species level.=E2=80=9D <o:p></o:p></p></div><div><p = class=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNormal>As I= recall, this was discussed but not acted upon until Layberry et al treated= all Canadian populations as A.stella in 1998. A year later, Opler doubled = down, adding a twist, treating A. thoosa as a distinct species as well. <o:= p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div= ><p class=3DMsoNormal>These publications created difficulties as I prepared= the catalogue but I had a friend (TS) who had published a paper in 2005 tr= eating A.sara and A.stella as distinct species in a rearing study. We talke= d and he said (something like): JP...I=E2=80=99m crazy for orange-tips and = I=E2=80=99m going to figure this one out - but I haven=E2=80=99t yet.=E2=80= =9D <o:p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></d= iv><div><p class=3DMsoNormal>I deferred from accepting the split in 2008. I= eagerly awaited Todd=E2=80=99s paper...which arrived in 2018 and I am grat= ified that Opler et al, 2022 supported Todd=E2=80=99s conclusions <o:p></o:= p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div><p cl= ass=3DMsoNormal>This took 22 years but I am not citing this as any sort of = standard! It is only an example. <o:p></o:p></p></div><div><p class=3DMsoNo= rmal><o:p> </o:p></p></div><div><p class=3DMsoNormal>JPPelham <o:p></o= :p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><div><p c= lass=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNormal><stro= ng><span style=3D'font-family:"Calibri",sans-serif'>Geiger, Hansj=C3=BCrg a= nd Arthur Maurice Shapiro</span></strong> <o:p></o:p></p></div><div><p clas= s=3DMsoNormal>1986 Electrophoretic evidence for speciation within the nomin= al species <em><span style=3D'font-family:"Calibri",sans-serif'>Anthocharis= sara</span></em> Lucas (Pieridae). <em><span style=3D'font-family:"Calibri= ",sans-serif'>Journal of Research on the Lepidoptera 25(1): 15-24, 2 figs.,= 4 tbls. {Spring 1986, 1 Oct 1986}</span></em> <o:p></o:p></p></div><div><p= class=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNormal><st= rong><span style=3D'font-family:"Calibri",sans-serif'>Layberry, Ross A., Pe= ter W. Hall and James Donald Lafontaine</span></strong> (plates by John T. = Fowler) <o:p></o:p></p></div><div><p class=3DMsoNormal>1998 <em><span style= =3D'font-family:"Calibri",sans-serif'>The butterflies of Canada</span></em>= . Toronto, Buffalo & London; University of Toronto Press: 280 pp., 32 p= ls., 294 maps and ill. {1998} <o:p></o:p></p></div><div><p class=3DMsoNorma= l><o:p> </o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p= ></div><div><p class=3DMsoNormal><strong><span style=3D'font-family:"Calibr= i",sans-serif'>Opler, Paul Alexander</span></strong> <o:p></o:p></p></div><= div><p class=3DMsoNormal>1999 <em><span style=3D'font-family:"Calibri",sans= -serif'>A field guide to western butterflies</span></em>. Boston, New York;= Houghton Mifflin Company, The Peterson Field Guide Series (Amy Bartlett Wr= ight (illustrator): xiv, [2], 540 pp., 44 pls., figs. & maps. {1999} <o= :p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><di= v><p class=3DMsoNormal><strong><span style=3D'font-family:"Calibri",sans-se= rif'>Stout, Todd L.</span></strong> <o:p></o:p></p></div><div><p class=3DMs= oNormal>2005 The challenge of raising northern Utah orangetips (<em><span s= tyle=3D'font-family:"Calibri",sans-serif'>Anthocharis stella browningi</spa= n></em> and <em><span style=3D'font-family:"Calibri",sans-serif'>Anthochari= s sara thoosa</span></em>). <em><span style=3D'font-family:"Calibri",sans-s= erif'>Bulletin of the Utah Lepidopterists=E2=80=99 Society 12(1): 5-9, 13, = 7 figs. {Mar 2005}</span></em> <o:p></o:p></p></div><div><p class=3DMsoNorm= al><span style=3D'font-size:12.0pt;font-family:Helvetica;color:#333333'><o:= p> </o:p></span></p></div><div><p class=3DMsoNormal><strong><span styl= e=3D'font-family:"Calibri",sans-serif'>Scott, James Allen and Michael S. Fi= sher</span></strong> <o:p></o:p></p></div><div><p class=3DMsoNormal>2008 <e= m><span style=3D'font-family:"Calibri",sans-serif'>Anthocharis</span></em> = =E2=80=9C<em><span style=3D'font-family:"Calibri",sans-serif'>sara</span></= em>=E2=80=9D group, especially in Colorado and vicinity (with new research = from Todd Stout) (one ssp. coauthored by Norbert G. Kondla). Pp. 1-14, pl. = 2, in: J. Scott (Ed.), Geographic variation and new taxa of western North A= merican butterflies, especially from Colorado. <em><span style=3D'font-fami= ly:"Calibri",sans-serif'>Papilio (New Series) 18 {3 Dec 2008}</span></em> <= o:p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><d= iv><p class=3DMsoNormal><strong><span style=3D'font-family:"Calibri",sans-s= erif'>Stout, Todd L.</span></strong> <o:p></o:p></p></div><div><p class=3DM= soNormal>2010 Observations on <em><span style=3D'font-family:"Calibri",sans= -serif'>Anthocharis julia browningi</span></em> and <em><span style=3D'font= -family:"Calibri",sans-serif'>Anthocharis thoosa thoosa</span></em> includi= ng tension zones near Nephi, Juab County, Utah. <em><span style=3D'font-fam= ily:"Calibri",sans-serif'>Taxonomic Report of the International Lepidoptera= Survey 7(4): 1-12 {31 May 2010}</span></em> <o:p></o:p></p></div><div><p c= lass=3DMsoNormal><o:p> </o:p></p></div><div><p class=3DMsoNormal><stro= ng><span style=3D'font-family:"Calibri",sans-serif'>Stout, Todd L.</span></= strong> <o:p></o:p></p></div><div><p class=3DMsoNormal>2018 A review of thr= ee species-level taxa of the <em><span style=3D'font-family:"Calibri",sans-= serif'>Anthocharis sara</span></em> complex (Lepidoptera: Pieridae: Pierina= e: Anthocharidini). <em><span style=3D'font-family:"Calibri",sans-serif'>In= secta Mundi 0615: 1-38, 26 figs., 3 tbls.,1 app. {30 Mar 2018}</span></em> = <o:p></o:p></p></div><div><p class=3DMsoNormal><o:p> </o:p></p></div><= div><p class=3DMsoNormal><strong><span style=3D'font-family:"Calibri",sans-= serif'>Opler, Paul Alexander, Todd L. Stout, Werner Back, Jing Zhang, Qian = Cong. Jinhui Shen, Jing Zhang and Nicolay Vyacheslav Grishin</span></strong= > <o:p></o:p></p></div><div><p class=3DMsoNormal>2022 DNA barcodes reveal d= ifferent speciation scenarios in the four North American <em><span style=3D= 'font-family:"Calibri",sans-serif'>Anthocharis</span></em> Boisduval, Rambu= r, [Dum=C3=A9nil] & Graslin, [1833] (Lepidoptera: Pieridae: Pierinae: A= nthocharidini) species groups. <em><span style=3D'font-family:"Calibri",san= s-serif'>Zootaxa 5194(4): 519=E2=80=93539, 28 figs. {7 Oct 2022}</span></em= > <o:p></o:p></p></div><div><p class=3DMsoNormal></o:p></span></p></div></d= iv></body></html> <div width=3D"1" style=3D"color:white;clear:both">_._,_._,_</div> <hr> Groups.io Links:<p> =20 You receive all messages sent to this group. =20 =20 <p> <a target=3D"_blank" href=3D"https://groups.io/g/NorWestLeps/message/11202"= >View/Reply Online (#11202)</a> | <a target=3D"_blank" href=3D"mailto:<NorWestLeps...>?subject=3DRe:%2= 0Re%3A%20%5BNorWestLeps%5D%20Taxonomic%20change%20-%20Anthocharis%20sara%20= complex">Reply To Group</a> =20 | <a target=3D"_blank" href=3D"mailto:<nel-de-bearck...>?subject= =3DPrivate:%20Re:%20Re%3A%20%5BNorWestLeps%5D%20Taxonomic%20change%20-%20An= thocharis%20sara%20complex">Reply To Sender</a> =20 | <a target=3D"_blank" href=3D"https://groups.io/mt/96507103/795800">Mute T= his Topic</a> | <a href=3D"https://groups.io/g/NorWestLeps/post">New Topic</a><br> <a href=3D"https://groups.io/g/NorWestLeps/editsub/795800">Your Subscriptio= n</a> | <a href=3D"mailto:NorWestLeps+<owner...>">Contact Group Owner</a> | <a href=3D"https://groups.io/g/NorWestLeps/leave/4559582/795800/778100922/x= yzzy">Unsubscribe</a> [<lists...>]<br> <div width=3D"1" style=3D"color:white;clear:both">_._,_._,_</div> --B_3757414716_2384768163-- |
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